The Kremling Kuthroats was founded by the evil old Kritters all over Crocodile Isle, after the end of the ancient Kremling society that was kept on Lost World. They started to establish a town above the second cliff, they built galleons, stole various values where they kept them in chests and deposit them in caves on the island itself. The oldest members were Kackle, Kloak and other Kritters with the pirate attire.
The Kremling Kuthroats was reorganized into the Kremling Krew by K. Rool who was a veteran pirate too.
Donkey Kong franchise
In Donkey Kong Country 2: Diddy's Kong Quest, veteran pirate Kremlings from Kremling Kuthroats as Kloak and Kackle are seen as ghosts living in the old town that was dilapidated during the Kremling Krew's reign. In that same game as the alive pirate veteran K. Rool orders the Kremling Krew members to return to their origins wearing the pirate attire from Kremling Kuthroats era. They are even feared by members of the Kremling Krew for being undeads.
^ abDonkey Kong Country 2: Diddy's Kong Quest Player's Guide, Nintendo, 1995, p. 20 - "GLOOMY GULCH: Don't count on meeting another living soul in this dilapidated ghost town, where even the ropes fade in and out of existence like phantoms."
^ abNintendo Power, Vol. 79, December, 1995, page 26 - "GLOOMY GULCH: The last stretch of road leading to K. Rool's Keep passes though the ghost town of Gloomy Gulch, and you'll be up to your hips in ghosties, beasties and bogles."
^Nintendo Magazine System (AU), Vol. 36, March, 1996, p. 21 - "TREASURE CHEST: Like all good pirates the Kremlings stash their swag in Treasure Chests."
^ abDonkey Kong 64 Player's Guide, Nintendo, 1999, p. 66 - "THE PIRATE'S HOARD: Transform into Enguarde and leap out of the water and through the star three times to open the way into the pirate's treasure room."
^ abNintendo Power, Vol. 76, September, 1995, page 26 "CAPTURED: Keep a watch out for Klobber when he's barreling ahead and the Ghost Pirate who haunts the forest shadows."
^M. Arakawa. Donkey Kong Country 2: Diddy's Kong Quest Player's Guide. p. 20 - "KREM QUAY: K. Rool proves he’s pirate and not a sailor with yet another wrecked ship. Be careful or you’ll get sunk in this swamp along with it!"
^Nintendo Power, Vol. 79, December, 1995, p. 22 - "KREM QUAY: What? Another wrecked ship? Hasn’t K. Rool ever learned how to drive these things?!"
^Donkey Kong Country 2: Diddy's Kong Quest Instruction Booklet, Nintendo, 1995, p. 30 - "KACKLE & KLOAK: These ghostly, Kremling spirits haunt many locations on Crocodile Isle."
^Nintendo Power, Vol. 76, September, 1995, p. 26 - "CAPTURED: They're the same, but not the same. They're deranged, estranged, and subtly changed. K. Rool's Kremling crew show its true colors in DKC2. No longer are they merely repulsive reptiles."
^Nintendo Power, Vol. 79, December, 1995, p. 15 - "CROCODILE ISLE: King K. Rool now fancies himself a pirate, taking on the title of "Kaptain" and ordering his Kremlings to dress up as peg-legged buccaneers."
^Donkey Kong Country 2: Diddy's Kong Quest German Player's Guide. p. 84 - "WEB WOODS:Dieser schrecklich düstere Wald wird sogar von den Kremlingpiraten gemieden."
^Donkey Kong Country 2: Diddy's Kong Quest Instruction Booklet, Nintendo, 1995, p. 22 - "GLOOMY GULCH: This spooky area is also haunted by the evil Kremling spirits known as Kloak and Kackle."
^"Mermaid:The reptiles have stolen my precious pearls and locked them inside a giant treasure chest!" - Donkey Kong 64